Spring vs Winter Alex E 6:59:00 AM Nature Edit Spring road. Andrew Nuding. Early spring. Early spring. Spring. Forest road. Spring. Bench in the park. Spring in the Russian city. Russian courtyard in the city of Stavropol. Share on Facebook Share on Twitter Share on Google Plus Alex E Ecoclimax is defined by Odum (1969) as the culmination state after a succession in a stabilized ecosystem in which maximum biomass (or high information content) and symbiotic function among organisms is kept per unit of available energy flow. RELATED POSTS Please enable JavaScript to view the comments powered by Disqus.