Humphead Parrotfish are rather extraordinary creatures: with their ultra-strong jaws, they eat coral reefs (and rock) and defecate sand — which in turn becomes the tropical beaches we know and love.
 
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Alex E

Ecoclimax is defined by Odum (1969) as the culmination state after a succession in a stabilized ecosystem in which maximum biomass (or high information content) and symbiotic function among organisms is kept per unit of available energy flow.